1. Research Institute for the Environment and Livelihoods, Charles Darwin University, Darwin, Northern Territory 0909, Australia
2. CSIRO Land and Water, Tropical Ecosystems Research Centre, Winnellie, Northern Territory 0822, Australia
3. School of Environmental Sciences, University of Liverpool, Liverpool L69 3GP, UK
4. School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Wits 2050, South Africa