1. Department of Biology, University of Utah, 257 South, 1400 East, Salt Lake City, UT, 84112–0840, USA.
2. Entomology Section, Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, CA 90007, USA.
3. Department of Biology, Universiti Brunei Darussalam, Bandar Seri Begawan, Brunei Darussalam.