1. Okazaki Institute for Integrative Bioscience, National Institute for Basic Biology (NIBB), National Institutes of Natural Sciences, 5-1 Myodaiji-Higashiyama, Okazaki 444-8787, Japan.
2. Graduate School of Biological Sciences, Nara Institute of Science and Technology, 8916-5 Takayama, Ikoma, Nara 630-0192, Japan.
3. Université de Toulouse, Université Paul Sabatier, Centre de Biologie du Développement, Bâtiment 4R3, 118 route de Narbonne, F-31062 Toulouse, France.
4. CNRS, UMR 5547, Centre de Biologie du Développement (CBD), F-31062 Toulouse, France.
5. Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency (JST), 4-1-8 Honcho, Kawaguchi, Saitama 332-0012, Japan.
6. Department of Basic Biology, School of Life Science, Graduate University for Advanced Studies (SOKENDAI), 38 Myodaiji-Nishigonaka, Okazaki 444-8585, Japan.