1. Center for Animal Resources and Development (CARD) and Graduate School of Molecular and Genomic Pharmacy, Kumamoto University, Honjo 2-2-1, Kumamoto 860-0811, Japan
2. Program in Developmental Biology, The Hospital for Sick Children, and Department of Molecular and Medical Genetics, University of Toronto, Ontario M5G 1X8, Canada
3. Department of Molecular Neuropathology, Brain Science Institute, The Institute of Physical and Chemical Research (RIKEN), 2-1 Hirosawa, Wako, Saitama, 351-0198, Japan
4. Mammalian Genetics laboratory, National Institute of Genetics, Mishima, Japan
5. Western Regional Research Center, ARS, USDA, 800 Buchanan Street, Albany CA 94710, USA