1. Marine Biological Association of the UK, The Laboratory 1 , Citadel Hill, Plymouth PL1 2PB , United Kingdom
2. School of Biological and Marine Sciences, University of Plymouth 2 , Plymouth, PL4 8AA , UK
3. University of Vienna 3 Department of Neurosciences and Developmental Biology, Faculty of Life Sciences , , Vienna, 1030 , Austria
4. Doctoral School of Ecology and Evolution, University of Vienna 4 , Vienna, 1030 , Austria