1. Department of Biology, Institute of Zoology, University of Fribourg, Switzerland
2. Department of Biology, Institute of Cell Biology, University of Bern, Switzerland
3. These authors contributed equally to this work
4. Department of Biology, Interfaculty Bioinformatics Unit, University of Bern, Switzerland
5. UPMC-CNRS FR2424, Station Biologique de Roscoff, Roscoff, France
6. Departament de Genètica, Universitat de Barcelona, Catalonia, Spain
7. Institut Català de Recerca i Estudis Avancats (ICREA), Barcelona, Spain
8. Institute of Zoology and Center of Molecular Bioscience Innsbruck, University of Innsbruck, Austria