1. Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences, Tianjin 300308, People's Republic of China
2. Metabolic and Biomolecular Engineering National Research Laboratory, Korea Advanced Institute of Science and Technology, Yuseong-gu, Daejeon 34141, Republic of Korea
3. College of Life Sciences, University of Chinese Academy of Sciences, Beijing 100049, People's Republic of China