1. Laboratory for Cell Recovery Mechanisms, Advanced Technology Development Center, RIKEN Brain Science Institute, Wako, Saitama 351-0198, Japan
2. Laboratories for Cell Biology and Neuroscience, Graduate School of Medicine, Osaka University, Suita, Osaka 565-0871, Japan
3. Laboratory for Cell Function and Dynamics, Advanced Technology Development Center, RIKEN Brain Science Institute, Wako, Saitama 351-0198, Japan
4. Structure and Function of Biomolecules, Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Corporation (JST), Nittochi 535, Akinono-cho, Nakagyo-ku, Kyoto 604-0847, Japan