1. Key Laboratory of Forest Genetics and Biotechnology Ministry of Education, Nanjing Forestry University, Nanjing, Jiangsu, China;
2. Co-Innovation Center for the Sustainable Forestry in Southern China, Nanjing, Jiangsu, China;
3. College of Biology and the Environment, Nanjing Forestry University, Nanjing, Jiangsu, China;
4. Experimental Center of Desert Forestry, Chinese Academy of Forestry, Dengkou, Inner Mongolia, China;
5. The Center for Comparative Oncology, University of California at Davis, Davis, CA, USA