1. Council for Scientific and Industrial Research (CSIR), Biosciences, Pretoria, South Africa
2. Department of Consumer and Food Sciences, University of Pretoria, Pretoria, South Africa
3. Centre de coopération internationale en recherche agronomique pour le développement (CIRAD), UMR – Qualisud, Montpellier Cedex 5, France
4. Department of Biochemistry, Genetics and Microbiology, University of Pretoria, Pretoria, South Africa