1. Department of Biological Chemistry, John Innes Centre, Norwich NR4 7UH, United Kingdom
2. School of Biological Sciences, University of East Anglia, Norwich NR4 7TJ, United Kingdom
3. Norwich Medical School, University of East Anglia, Norwich NR4 7UQ, United Kingdom
4. Department of Crop Genetics, John Innes Centre, Norwich NR4 7UH, United Kingdom