1. Department of Ecology and Evolution, University of Lausanne, 1015 Lausanne, Switzerland (P.-A.C., N.S., G.B.); Swiss Institute of Bioinformatics, 1015 Lausanne, Switzerland (N.S.); Department of Biology, University of Missouri, St. Louis, Missouri 63121 (E.A.K.); and Center for Tropical Forest Science, Smithsonian Tropical Research Institute, Apartado Postal 0843–03092, Panama, Republic of Panama