1. Forage Improvement Division (C.Z., L.H., C.F., Q.J., J.H.B., Z.-Y.W.) and Plant Biology Division (C.P., J.N., L.Q., E.B.B., Y.T., M.U.), Samuel Roberts Noble Foundation, Ardmore, Oklahoma 73401; Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, School of Life Sciences, Shandong University, Jinan 250100, China (L.H., G.X.)