1. Department of Wheat and Barley Research, National Institute of Crop Science, National Agricultural Research Organization, 2–1–18 Kannondai, Tsukuba, Ibaraki 305–8518, Japan (M.C., I.H., T.H., Y.W.); Department of Applied Biochemistry (H.Z.) and Center for Research on Wild Plants (K.Y.), Utsunomiya University, 350 Mine-machi, Utsunomiya, Tochigi 321–8505, Japan; Barley Germplasm Center, Research I