1. Warwick HRI, Warwick CV35 9EF, United Kingdom (C.R.H., H.C.B., J.P.H., A.M., K.A.P., P.J.W.); School of Biological Sciences, The University of Birmingham, Birmingham B15 2TT, United Kingdom (C.R.H., J.P.); and Plant Sciences Division, University of Nottingham, Sutton Bonington Campus, Loughborough LE12 5RD, United Kingdom (M.R.B., J.P.H., K.A.P.)