1. Faculty of Agriculture, Kyushu University, Fukuoka 812-8581, Japan
2. Institute of Molecular, Cell, and Systems Biology, College of Medical, Veterinary, and Life Sciences, University of Glasgow, Glasgow G12 8QQ, United Kingdom
3. Graduate School of Biostudies, Kyoto University, Kyoto 606-8502, Japan
4. Graduate School of Science, University of Tokyo, Tokyo 113-0033, Japan
5. Graduate School of Science and Engineering, Tokyo Metropolitan University, Tokyo 192-0397, Japan