1. Institute of Biological Sciences, University of Tsukuba, Tsukuba, Ibaraki, 305–8572, Japan (M.A., H.I., A.K., H.K., S.S.); and
2. Plant Science Center, The Institute of Physical and Chemical Research (RIKEN), Wako, Saitama, 351–0198, Japan (S.Y., Y.K.)