1. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China
2. College of Life Sciences Gannan Normal University Ganzhou 34100 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Hunan Engineering Technology Research Center of Veterinary Drugs Hunan Agricultural University Changsha 410128 China
5. Department of Ecology and Evolutionary Biology University of Toronto 25 Willcocks St. Toronto ON M5S 3B2 Canada