1. Department of Life Sciences, University of Dundee, Dundee DD1 4HN, Scotland, United Kingdom (J.M.L, A.J.F.); John Innes Centre, Norwich Research Park, Colney, Norwich NR4 7UH, United Kingdom (G.F.D., T.H.N.E., M.J.A., J.D.); Scottish Crop Research Institute, Invergowrie DD2 5DA, Scotland, United Kingdom (D.M.); and Centre for Genetic Resources, 6700AA Wageningen, The Netherlands (T.J.L.v.H.)