1. State Key Laboratory of Grassland Agro-ecosystems; Key Laboratory of Grassland Livestock Industry Innovation, Ministry of Agriculture and Rural Affairs; College of Pastoral Agriculture Science and Technology, Lanzhou University, Lanzhou 730000, People’s Republic of China
2. Ministry of Education Key Laboratory of Cell Activities and Stress Adaptations, School of Life Sciences, Lanzhou University, Lanzhou 730000, People’s Republic of China
3. Department of Biology, Institute of Biochemistry, Carleton University, Ottawa, Ontario, K1S 5B6, Canada