1. Graduate School of Agricultural and Life Sciences, The University of Tokyo, Japan
2. GPS Co. LTD., Saitama, Japan
3. Hokkaido Regional Breeding Office, Forest Tree Breeding Center, Forestry and Forest Products Research Institute, Hokkaido, Japan Present address
4. Section of Nature and Environment, Institute of Natural and Environmental Sciences, University of Hyogo, Japan
5. Laboratory of Conservation Ecology, Department of Integrated Science and Engineering for Sustainable Society, Faculty of Science and Engineering, Chuo University, Tokyo, Japan