1. Hebei Provincial Key Laboratory of Forest Trees Germplasm Resources and Forest Protection, and College of Forestry, Agricultural University of Hebei, Lekai South Street 2596, Baoding 071000, China
2. Key Laboratory of Forest Ecology and Environment of the State Administration of Forestry, and Institute of Forest Ecology, Environment and Protection, Chinese Academy of Forestry, Beijing 100091, China
3. Faculty of Forestry, University of British Columbia, Forest Sciences Center, 2211–2424 Main Mall, Vancouver, BC, V6T1Z4, Canada
4. Co-Innovation Center for Sustainable Forestry in Southern China, Nanjing Forestry University, Nanjing, Jiangsu 210000, China