1. Guangxi Key Laboratory of Karst Ecological Processes and Services, Institute of Subtropical Agriculture, Chinese Academy of Sciences, Changsha, China
2. Huanjiang Observation and Research Station for Karst Ecosystem, Chinese Academy of Sciences, Huanjiang, China
3. University of Chinese Academy of Sciences, Beijng, China
4. Department of Geosciences and Natural Resource Management, University of Copenhagen, Copenhagen, Denmark
5. Department of Geography and Geology, Algoma University, Canada