1. The Fresh Produce Research Centre, Crop and Environment Sciences Department, Harper Adams University, Edgmond, Shropshire, UK
2. Warwick Crop Centre, University of Warwick, Wellesbourne, Warwickshire, UK
3. Department of Biology, University of York, Heslington, York, UK
4. Functional and Comparative Genomics, Institute of Integrative Biology, University of Liverpool, Liverpool, UK