1. ICORP, Computational Brain Project, Japan Science and Technology Agency, Saitama 332-0012, Japan, ATR Computational Neuroscience Laboratories, Department of Humanoid Robotics and Computational Neuroscience, Kyoto 619-0288, Japan,
2. Computer Science and Artificial Intelligence Laboratory, Massachusetts Institute of Technology, Cambridge, MA 02139, USA,
3. Computer Science & Neuroscience, University of Southern California, Los Angeles, CA 90089-2520, USA,
4. ATR Computational Neuroscience Laboratories, Department of Humanoid Robotics and Computational Neuroscience, Kyoto 619-0288, Japan, Computer Science & Neuroscience, University of Southern California, Los Angeles, CA 90089-2520, USA,