1. Palaeoecology, Institute of Plant Sciences, University of Bern, Altenbergrain 21, CH-3013 Bern, Switzerland,
2. Palaeoecology, Laboratory of Palaeobotany and Palynology, Institute of Environmental Biology, Utrecht University, Budapestlaan 4, 3584 CD Utrecht, The Netherlands
3. Palaeoecology, Institute of Plant Sciences, University of Bern, Altenbergrain 21, CH-3013 Bern, Switzerland, Palaeoecology and Vegetation Dynamics, Institute of Terrestrial Ecosystems, Department of Environmental Sciences, ETH Zurich, Universitätstrasse 16, CH-8092 Zurich, Switzerland