1. *State Key Laboratory of Agricultural Microbiology; Engineering Research Center of Green Development for Conventional Aquatic Biological Industry in the Yangtze River Economic Belt, Ministry of Education; College of Fisheries, Huazhong Agricultural University, Wuhan, China;
2. †School of Marine Science and Engineering, Qingdao Agricultural University, Qingdao, China;
3. ‡Hubei Hongshan Laboratory, Wuhan, China; and
4. §Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China