1. *State Key Laboratory of Bioreactor Engineering, East China University of Science and Technology, Shanghai, China;
2. †Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai, China;
3. ‡Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao, China; and
4. §Shanghai Engineering Research Center of Maricultured Animal Vaccines, Shanghai, China