1. School of Life Science, Liaoning Normal University, Dalian 116029, China
2. Liaoning Key Laboratory of Marine Animal Immunology and Disease Control, Dalian Ocean University, Dalian 116023, China
3. Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai 519000, China
4. Laboratory of Marine Fisheries Science and Food Production Process, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266235, China
5. Dalian Key Laboratory of Aquatic Animal Disease Prevention and Control, Dalian Ocean University, Dalian 116023, China