1. Zhejiang University, Department of Marine Sciences, Zhoushan, China.
2. Zhejiang University, Department of Marine Sciences, Zhoushan, China and Zhejiang University, Hainan Institute, Sanya, China. (corresponding author)
3. Zhejiang University, Department of Marine Sciences, Zhoushan, China and Zhejiang University, Hainan Institute, Sanya, China.