1. Graduate School of Engineering and Science, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan; Email: acheilognathus5884@gmail.com. Send reprint requests to this address.
2. Faculty of Fisheries and Marine Science, Sam Ratulangi University, Manado 95115, Indonesia; Email: sabanib@yahoo.com.
3. Tropical Biosphere Research Center, University of the Ryukyus, 1 Senbaru, Nishihara, Okinawa 903-0213, Japan; Email: (KY) yamahira@lab.u-ryukyu.ac.jp.