1. Faculty of Fisheries and Marine Science, Halu Oleo University, Kendari 93232, Indonesia
2. Faculty of Fisheries and Marine Science, Sam Ratulangi University, Manado 95115, Indonesia
3. Department of Environmental Science, Fukuoka Women's University, Fukuoka 813-8529, Japan
4. Department of Biology, Osaka Medical and Pharmaceutical University, Osaka 569-8686, Japan
5. Ecological Genetics Laboratory, National Institute of Genetics, Mishima, Japan
6. Tropical Biosphere Research Center, University of the Ryukyus, Okinawa 903-0213, Japan