1. Department of Biological Sciences, University of Calgary, Calgary, Canada
2. McCaig Institute for Bone and Joint Health, Calgary, Canada
3. Department of Cell Biology and Anatomy, University of Calgary, Calgary, Canada
4. Alberta Children's Hospital Research Institute for Child and Maternal Health, University of Calgary, Calgary, Canada
5. Department of Comparative Biology and Experimental Medicine, Faculty of Veterinary Medicine, University of Calgary, Calgary, Canada