1. The University of Tokyo,International Research Center for Neurointelligence (WPI-IRCN),Bunkyo City, Tokyo,Japan,113-0033
2. Graduate School of Medicine, Osaka University, and Japan Society for the Promotion of Science,Department of Advanced Hybrid Medicine,Tokyo,Japan
3. Graduate School of Education, Kyoto University,Tokyo,Japan
4. The University of Tokyo,Research Center for Neurointelligence (WPI-IRCN),Tokyo,Japan