1. Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington, 10th and Constitution Ave., Washington, D.C. 20560-0166, U.S.A.;, Email: bortedwards@gmail.com
2. Department of Biology, San Francisco State University, 1600 Holloway Ave., San Francisco, California 94132, U.S.A.
3. Department of Botany, University of Hawai'i, Mānoa, 3190 Maile Way, Honolulu, Hawaii 96822, U.S.A.
4. Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington, 10th and Constitution Ave., Washington, D.C. 20560-0166, U.S.A.