1. INATAA, Université Constantine 1 Frères Mentouri, Route de Ain El Bey, Constantine 25000, Algeria
2. Laboratory of natural substances, biomolecules and biotechnological applications, Faculty of Nature and Life Sciences, Université Oum El Bouaghi, Algeria
3. PEGASE, INRAE, Institut Agro, 35590 Saint-Gilles, France