1. Research Center for Plant Conservation, Botanic Gardens and Forestry, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor Km. 46, Bogor, Jawa Barat, 16911, Indonesia
2. Research Center for Food Crops, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor Km. 46, Bogor, Jawa Barat, 16911, Indonesia
3. Division of Applied Life Science, Graduate School of Agriculture, Kyoto University, Kyoto, 606-8502, Japan