1. Department of Animal Production, Escola Superior de Agricultura “Luiz de Queiroz,” University of Sao Paulo (ESALQ-USP), Piracicaba, SP, Brazil, 13418-900
2. Embrapa Beef Cattle, Campo Grande, MS, Brazil
3. Department of Exact Science, ESALQ-USP, Piracicaba, SP, Brazil
4. Department of Animal Science, University of California, Davis 95616