1. Division of Animal & Dairy Science, Chungnam National University, Dajeon 34134, South Korea
2. Hanwoo Research Institute, National Institute of Animal Science, PyeongChang 25340, South Korea
3. Division of Animal Production, National Institute of Animal Science, Wanju 55365, South Korea
4. Animal Genetic Improvement Division, National Institute of Animal Science, Seonghwan 31000, South Korea