1. Cooperative Research Centre (CRC) for Beef Genetic Technologies, University of New England, Armidale, New South Wales 2351, Australia
2. Commonwealth Scientific and Industrial Research Organisation (CSIRO) Livestock Industries, Queensland Bioscience Precinct, Brisbane, Queensland 4067, Australia
3. Animal Genetics and Breeding Unit, University of New England, Armidale, New South Wales 2351, Australia
4. The University of Queensland, School of Veterinary Science, Queensland 4072, Australia
5. Department of Employment, Economic Development and Innovation (DEEDI), Queensland 4820, Australia