1. Institute of Zoology, Zoological Society of London, Regent's Park, London NW1 4RY, United Kingdom
2. Department of Wildlife Ecology and Conservation, University of Florida, Gainesville, Florida 32611 USA
3. Instituto de Ecología, A.C., Av. Lázaro Cárdenas No. 253, A.P. 386, C.P. 61600 Pátzcuaro, Michoacán, México
4. School of Graduate Research, University of Waikato, Private Bag 3105, Hamilton 3240, New Zealand
5. ARC Centre of Excellence for Coral Reef Studies, James Cook University, Townsville, QLD, 4811, Australia