1. Graduate School of Science and Technology, Keio University, Yokohama, Kanagawa, 223-8522, Japan
2. Swartz Center for Computational Neuroscience, Institute for Neural Computation, University of California, San Diego, La Jolla, California, 92093, USA
3. Swartz Center for Computational Neuroscience, Institute for Neural Computation, Center for Advanced Neurological Engineering, Institute of Engineering in Medicine, University of California, San Diego, La Jolla, California, 92093, USA