1. Savannah River Ecology Laboratory, University of Georgia, PO Drawer E, Aiken, South Carolina, USA
2. Daniel B. Warnell School of Forestry and Natural Resources, University of Georgia, Athens, Georgia, USA
3. Archbold Biological Station, 123 Main Drive, Venus, Florida, USA
4. Department of Ecology and Evolutionary Biology, Cornell University, Ithaca, New York, USA