1. University of Hawai'i at Hilo, Department of Biology. 200 West Kāwili Street, Hilo, Hawai'i, 96720, United States of America
2. University of Hawai'i at Mānoa, Department of Oceanography, 1000 Pope Road, Honolulu, Hawai'i 96822, United States of America
3. Bishop Museum, Department of Natural Sciences 1525 Bernice St, Honolulu, Hawai‘i 96817, United States of America