1. Herbarium and Department of Evolution, Ecology, and Organismal Biology, The Ohio State University, 1315 Kinnear Road, Columbus, OH 43212, USA
2. Department of Botany and Biodiversity Research, University of Vienna, Rennweg 14, A-1030 Vienna, Austria
3. Department of Ecology and Evolutionary Biology and the Biodiversity Institute, University of Kansas, 1200 Sunnyside Avenue, Lawrence, KS 66045, USA