1. Department of Agricultural Sciences, Viikki Plant Science Centre, University of Helsinki , PO Box 27, Helsinki 00014, Finland
2. State Key Laboratory of Subtropical Silviculture, College of Forestry and Biotechnology, Zhejiang A&F University , Lin’an District, Hangzhou, Zhejiang 311300, China
3. Oak Ridge National Laboratory, Biosciences Division , 1 Bethel Valley Rd, Oak Ridge, TN 37830, USA
4. Production Systems Unit, Natural Resources Institute Finland (Luke) , Vipusenkuja 5, Savonlinna 57200, Finland
5. Institute of Biotechnology, University of Helsinki , PO Box 56, Helsinki 00014, Finland
6. Production Systems Unit, Natural Resources Institute Finland (Luke) , Paavo Havaksentie 3, Oulu 90570, Finland