1. Graduate School of Engineering Science, Akita University , Tegata Gakuen-machi 1-1, Akita City, Akita 010-0852 , Japan
2. Plant Breeding and Genetics Section, School of Integrative Plant Science, Cornell University , Ithaca, NY 14853 , USA
3. Institute of Biological Chemistry, Washington State University , Pullman, WA 99164 , USA
4. Research Institute for Sustainable Humanosphere, Kyoto University , Gokasho, Uji, Kyoto 611-0011 , Japan
5. School of Biological Sciences, The University of Hong Kong , Pokfulam, Hong Kong , China