1. Division of Life Science, Graduate School of Science and Engineering, Saitama University , Shimo-Okubo 255, Sakura-ku, Saitama, 338-8570 Japan
2. Metabolic Systems Research Team, RIKEN Center for Sustainable Resource Science , 1-7-22, Suehiro, Tsurumi, Yokohama, Kanagawa, 230-0045 Japan
3. Department of Biology, Tokyo Gakugei University , 4-1-1 Nukuikita-machi, Koganei, Tokyo, 184-8501 Japan
4. Department of Life Sciences, Graduate School of Arts and Sciences, The University of Tokyo , 3-8-1 Komaba, Meguro-ku, Tokyo, 153-8902 Japan
5. Department of Applied Biosciences, Graduate School of Bioagricultural Sciences, Nagoya University , Furo-cho, Chikusa-ku, Nagoya, 464-8601 Japan