1. Key Laboratory of Animal Genetics, Breeding and Reproduction in the Plateau Mountainous Region, Ministry of Education, College of Animal Science, Guizhou University, Guiyang 550025, China
2. College of Grassland Agriculture, Northwest A & F University, Yangling 712100, China
3. Rapeseed Research Institute, Guizhou Academy of Agricultural Sciences, Guiyang 550008, China
4. Grasslands and Sustainable Farming, Production Systems Unit, Natural Resources Institute Finland, Halolantie 31A, FI-71750 Kuopio, Finland