1. CAS Key Laboratory of Tropical Plant Resources and Sustainable Use, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences , Kunming, Yunnan 650223 , China
2. Center of Economic Botany, Core Botanical Gardens, Chinese Academy of Sciences , Kunming, Yunnan 650223 , China
3. University of Chinese Academy of Sciences , Beijing 100049 , China
4. School of Life Sciences, University of Science and Technology of China , Hefei, Anhui 230027 , China